Johann's pinyon

Pinus johannis is a small to medium-size tree, often just a shrub, reaching 4–10 metres (13–33 ft) tall and with a trunk diameter of up to 50 cm (20 in). The bark is grey-brown, thin and scaly at the base of the trunk. The leaves ('needles') are in mixed fascicles of three and four, slender, 3–6 cm (1+1⁄4–2+1⁄4 in) long, and deep green to blue-green, with stomata confined to a bright white band on the inner surfaces. The cones are globose, 2–4 cm (3⁄4–1+1⁄2 in) long and 2–3 cm (3⁄4–1+1⁄4 in) broad when closed, green at first, ripening yellow-brown when 16–18 months old, with only a small number of thin, fragile scales, typically 6-12 fertile scales. The cones open to 3–5 cm (1+1⁄4–2 in) broad when mature, holding the seeds on the scales after opening. The seeds are 9–12 mm (11⁄32–15⁄32 in) long, with a thick shell, a white endosperm, and a vestigial 1–2 mm (1⁄32–3⁄32 in) wing; they are dispersed by the Mexican jay, which plucks the seeds out of the open cones. The jay, which uses the seeds as a major food resource, stores many of the seeds for later use; some of these stored seeds are not used and are able to grow into new trees. Pinus johannis is nearly dioecious: nearly all individuals produce either male or female cones only, and trees do not appear to change their sex expression over time.

The edible pine nut seeds are collected in Mexico to a small extent. The white-glaucous inner surfaces of the needles make it a very attractive small tree, suitable for parks and large gardens.

Pinus johannis is a recently described pinyon pine, discovered by Elbert L. Little in 1968 when comparing pinyons growing in Arizona with those of typical Mexican pinyon (Pinus cembroides) in Mexico; he described it as a variety of Mexican pinyon, Pinus cembroides var. bicolor, noting the very different stomatal placing on the leaves; it also differs in needle number, with 3–4 per fascicle, rather than 2–3; in the cones having thinner scales; and in having a denser, more rounded crown. Further research by the French botanist Marie-Françoise Robert-Passini, the American botanists Dana K. Bailey and Frank G. Hawksworth and others, has shown that it is better treated as a distinct species. Although often occurring together with Mexican pinyon, it is reproductively isolated from that by its pollination being a month to two months later in summer, rather than in spring, thereby preventing hybridisation. As Robert-Passini and Bailey & Hawksworth were working in different areas at about the same time, it was raised to species rank twice, first as Pinus johannis by Robert-Passini (naming it after her husband Jean) examining specimens in the Sierra Madre Oriental in Mexico, and then later as Pinus discolor by Bailey & Hawksworth examining specimens in the northern Sierra Madre Occidental in Arizona. There are slight differences between the plants in the two ranges; those in the eastern …