Chelicerates

The subphylum Chelicerata (from Neo-Latin, from French chélicère, from Ancient Greek χηλή (khēlḗ) 'claw, chela' and κέρας (kéras) 'horn') constitutes one of the two major living subdivisions of the phylum Arthropoda, alongside Mandibulata. Chelicerates include the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids.

Chelicerata split from Mandibulata by the mid-Cambrian, as evidenced by stem-group chelicerates like Habeliida, Mollisonia and Megachelicerax present by this time. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species.