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Bombardier beetleNL·0316
Plate 1 of 4. (c) Francis Birlenbach, some rights reserved (CC BY-NC), uploaded by Francis Birlenbach · via iNaturalist

Herbarium of the Nature Library

Name
Brachinus crepitans
Family
Carabidae
Native to
Europe, especially the south and west, east into western Asia
Habitat
Calcareous grassland, Dry grassland, Coastal cliffs
IUCN NE · Not evaluated

Bombardier beetle

Brachinus crepitans

Also calledBombardierkäfer (German) · Brachyne pétard (French) · Escarabajo bombardero (Spanish)

A small ground beetle that defends itself with a boiling chemical explosion. Two harmless liquids stored in its abdomen are mixed with enzymes in a reaction chamber, firing a hot, caustic spray with an audible pop.

Brachinus crepitans is a species of ground beetle in the Brachininae subfamily that can be found in Europe, central and west Asia, and northern Africa. Wikipedia

§1

What it can do

  1. iMixes hydroquinones and hydrogen peroxide with catalase and peroxidase enzymes to produce a spray near 100 °C
  2. iiFires the spray in rapid pulses with an audible crack
  3. iiiCan aim the spray by swivelling the tip of its abdomen
§2

In the field

Field marks

  • aOrange-red head, thorax and legs
  • bDark metallic blue to blue-green wing cases
  • cSmall, narrow ground-beetle shape
  • dOften in groups under stones in dry grassland
Not to be confused with
Brachinus explodens is smaller (5–7.5 mm) and very similar; a hand lens and key are needed. Other red-and-blue ground beetles (e.g. Lebia, Drypta) lack the defensive pop.
Where to see it
Chalk downland and dry coastal grassland in southern England and continental Europe.
When
Adults mainly Apr–Sep
§3

Listen

Looking for field recordings…

§4

Life & habits

Diet
Adults are predators and scavengers of small invertebrates
Habitat
Warm, dry, sparsely vegetated ground, often on chalk and limestone; shelters under stones
Size
About 6–10 mm

When grabbed by an ant, spider or frog, it blasts the attacker with hot benzoquinones.

§5

Where it lives

Recorded sightings

—

geo-referenced records in GBIF, from museums, herbaria and field observers.

Native range

Europe, especially the south and west, east into western Asia

Drag the globe to turn it. Dots are where records exist, not a range map; darker means more records.

§6

In the stacks

Searching the stacks…

Every page naming Brachinus crepitans in the Biodiversity Heritage Library →

§7

Marginalia

Linnaeus named it crepitans, 'crackling', in 1758 for its explosive defence. In the late 20th century it became a famous example in debates between creationists and evolutionary biologists over how complex traits evolve.

Curiosity

High-speed X-ray imaging of bombardier beetles in 2015 showed that each spray is a rapid series of pulses, with an inlet valve to the reaction chamber closing at each blast.

§8

Kinship

  1. kingdomAnimalia
  2. ›phylumArthropoda
  3. ›subphylumHexapoda
  4. ›classInsecta
  5. ›subclassPterygota
  6. ›orderColeoptera
  7. ›suborderAdephaga
  8. ›superfamilyCaraboidea
  9. ›familyCarabidae
  10. ›subfamilyBrachininae
  11. ›tribeBrachinini
  12. ›genusBrachinus
  13. ›subgenusBrachinus
Open its branch on the Wheel of Life →

Photographs: iNaturalist contributors under the licences shown. Records: GBIF.org. Historical plates: Source Library (CC BY-SA 4.0; originals public domain). Summary text: Nature Library editors and Wikipedia (CC BY-SA). Medicinal and edibility notes are historical and educational, not advice. iNaturalist taxon 525781.