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NL·0162PlantsSolanaceaeHerb49,733 field observations
TomatoNL·0162
Plate 1 of 4. (c) alexkehl, some rights reserved (CC BY-NC) · via iNaturalist

Herbarium of the Nature Library

Name
Solanum lycopersicum
Family
Solanaceae
Native to
Wild ancestor (Solanum pimpinellifolium) in coastal Ecuador and Peru; domesticated in Mesoamerica; grown worldwide
Habitat
Dry coastal valleys (wild), Gardens and farmland
IUCN NE · Not evaluated

Tomato

Solanum lycopersicum

Also calledTamatar (Hindi/Urdu) · Xītomatl (Nahuatl) · Jitomate (Mexican Spanish) · Fānqié / Xīhóngshì (Chinese) · Banadura / Ṭamāṭim (Arabic) · Gojeh farangi (Persian) · Nyanya (Swahili)

A sprawling Andean nightshade, domesticated in Mexico, that Europeans long feared as poisonous before it conquered Italian and then world cuisine. Its pea-sized wild ancestor still grows in coastal Ecuador and Peru.

The tomato is a plant whose fruit is an edible berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco, potato, and chili peppers. It originated from western South America, and may have been domesticated there, in Mexico, or in Central America. The Spanish introduced tomatoes to Eurasia in the Columbian exchange in the 16th century. Wikipedia

§1

What it can do

  1. iFlowers are buzz-pollinated: bumblebees vibrate them to release pollen
  2. iiGlandular hairs on stems release the distinctive 'tomato' smell and trap small insects
  3. iiiFruit ripening is triggered by the plant hormone ethylene
  4. ivStems lying on soil sprout roots along their length
§2

At the table

Edible
Edible with care
Not eaten
Toxic
Deadly

Eatenfruit (raw, cooked, dried)

Ripe fruit is safe. Leaves and stems contain tomatine and other glycoalkaloids and are mildly toxic, especially to pets; green fruit has small amounts that are usually harmless when cooked.

§3

In the medicine cabinet

TraditionUse
Mesoamerican folk medicineFruit and leaves applied to skin infections and sore throats
Modern nutrition scienceLycopene from cooked tomatoes studied for prostate and cardiovascular health, with mixed results

Weight of evidence

Traditional. Long historical use; little or no modern clinical evidence.

Active compounds

lycopene, β-carotene, tomatine (leaves, green fruit), naringenin chalcone

CautionLeaves and stems are toxic to pets; tomatoes can worsen reflux.
§4

Fruit & seed

Fruit: Tomatoes Strawberry

Fruit

Berry

Ripens: Summer to autumn (year-round under cover)

Juicy berry 1–15 cm, round, oval, plum- or heart-shaped; red, yellow, orange, pink, green-striped or near-black; fleshy walls around gel-filled seed chambers.

Seed: Tomato seed stages2Seed: Tomato seed stages3Seed: Starr-110215-0963-Solanum lycopersicum-seed packet-KiHana Nursery Kihei-Maui (24956930012)

Seed

~3 mm

Flat, round to kidney-shaped, pale tan to buff seeds about 3 mm across, covered in fine silky hairs and coated in jelly in the fruit

Dispersal
Birds, tortoises (in the Galápagos) and mammals
Germination
Sow 0.5 cm deep at 20–28 °C; sprouts in 5–10 days. Ferment saved seed in its jelly for 2–3 days, then rinse and dry

Photographs from Wikimedia Commons under open licences; hover for the photographer, click for the file.

§5

How to grow it

Hardiness (USDA zones)

1
2
3
4
5
6
7
8
9
10
11
12
13

Annual warm-season crop; frost-tender perennial in USDA 10–12. Survives winters down to about -1 °C.

Light
Full sun (6–8+ hours)
Water
Moderate, steady; irregular watering causes splitting and blossom-end rot
Soil
Rich, well-drained loam, pH 6–7
Propagation
Seed; side-shoot cuttings root in water
To harvest
60–100 days from transplanting
Climates
Temperate (summer), Mediterranean, Subtropical, Tropical highland
  1. 1Sow indoors 6–8 weeks before the last frost; pot on seedlings when they have true leaves.
  2. 2After frost, harden off and plant deeply — bury the stem up to the first leaves, as it will root along its length.
  3. 3Stake or cage plants; on cordon (indeterminate) varieties, pinch out side shoots in leaf axils.
  4. 4Water deeply at the base and mulch to keep moisture even; feed with a high-potassium fertiliser once fruit sets.
  5. 5Pick when fully coloured; at season's end, ripen green fruit indoors in a bag with a banana.

Can it grow where you are?

or
§6

Where it lives

Recorded sightings

—

geo-referenced records in GBIF, from museums, herbaria and field observers.

Native range

Wild ancestor (Solanum pimpinellifolium) in coastal Ecuador and Peru; domesticated in Mesoamerica; grown worldwide

Drag the globe to turn it. Dots are where records exist, not a range map; darker means more records.

§7

In Gerard's Herball, 1597

Page 275 of Gerard's Herball, 1597: Of apples of Loue

Second Booke · Chap. 55 · p. 275 · read the page →

Of apples of Loue.

John Gerard, The Herball, or Generall Historie of Plantes, London, 1597. What England's best-known herbal said of tomato, in its own spelling.

The vertues
In Spaine and those hot regions they vse to eate the apples prepared and boiled with pepper, salt, and oile: but they yeelde very little nourishment to the bodie, and the same naught and corrupt. Likewise they do eate the apples with oile, vineger and pepper mixed together for sauce to their meate, euen as we in these cold countries do mustarde.
The place
Apples of Loue do growe in Spaine, Italie, and such hot countries, from whence my selfe haue receiued seedes for my garden, where they do increase and prosper.
The time
It is sowen in the beginning of Aprill in a bed of hot horse dung after the maner of muske Melons and such like colde fruits.
The temperature
The golden Apple with the whole herbe it selfe is colde, yet not fully so colde as Mandrake, after the opinion of Dodonaeus: but in my iudgement it is very colde, yea perhaps in the highest degree of coldnesse: my reason is, bicause I haue in the hottest time of sommer cut away the superfluous braun…
The names
The apple of Loue is called in Latine Pomum Aureum, Poma Amoris, and Lycopersium, of some Glaucium: in English apples of Loue, and golden apples: in French Pommes d'amours: howbeit there be other golden apples whereof the poets do fable growing in the gardens of the daughters of Hesperus, which a dragon was appointed to keepe, who, as they fable, was killed by Hercules.

Gerard wrote within Galenic humoral medicine, and his opinions, including about the peoples of the Americas and their crops, are those of 1590s London. His “vertues” are history, not advice. Transcription: Source Library (CC BY-SA 4.0). The whole chapter in the Herball Room.

§8

In the old herbals & bestiaries

Woodcuts and engravings matched from the Source Library’s scanned books by name. Plates marked “relative” show a close kin rather than this exact species. Click a plate to read the page it came from, with translation.

§9

In the stacks

Searching the stacks…

Every page naming Solanum lycopersicum in the Biodiversity Heritage Library →

§10

Marginalia

The Italian herbalist Pietro Andrea Mattioli called it 'pomi d'oro' (golden apples) in 1544, and Europeans grew it as an ornamental for decades fearing it was poisonous. The Spanish town of Buñol throws well over 100 tonnes of tomatoes each August at La Tomatina.

Curiosity

In 1893 the US Supreme Court ruled (Nix v. Hedden) that the tomato is legally a vegetable, though botanically a fruit.

§11

Kinship

  1. kingdomPlantae
  2. ›phylumTracheophyta
  3. ›subphylumAngiospermae
  4. ›classMagnoliopsida
  5. ›orderSolanales
  6. ›familySolanaceae
  7. ›subfamilySolanoideae
  8. ›tribeSolaneae
  9. ›genusSolanum
Open its branch on the Wheel of Life →

Fact-checked: 1 correction made to this sheet. Photographs: iNaturalist contributors under the licences shown. Records: GBIF.org. Historical plates: Source Library (CC BY-SA 4.0; originals public domain). Summary text: Nature Library editors and Wikipedia (CC BY-SA). Medicinal and edibility notes are historical and educational, not advice. iNaturalist taxon 51737.